Speciation does not necessarily promote evolutionary change; rather, speciation 'gathers in' and guards evolutionary change by locking and stabilization for sufficient geological time within a Darwinian individual of the appropriate scale. If a change in a local population does not gain such protection, it becomes-to borrow Dawkins's metaphor at a macroevolutionary scale-a transient duststorm in the desert of time, a passing cloud without borders, integrity, or even the capacity to act as a unit of selection, in the panorama of life's phylogeny.
Life shows no trend to complexity in the usual sense-only an asymmetrical expansion of diversity around a starting point constrained to be simple.
The history of life is a tale of decimation and later stabilization of few surviving anatomies, not a story of steady expansion and progress.
Can I pay any higher tribute to a man [George Gaylord Simpson] than to state that his work both established a profession and sowed the seeds for its own revision? If Simpson had reached final truth, he either would have been a priest or would have chosen a dull profession. The history of life cannot be a dull profession.
Life is a copiously branching bush, continually pruned by the grim reaper of extinction, not a ladder of predictable progress.
Among people I have met, the few whom I would term “great” all share a kind of unquestioned, fierce dedication; an utter lack of doubt about the value of their activities (or at least an internal impulse that drives through any such angst); and above all, a capacity to work (or at least to be mentally alert for unexpected insights) at every available moment of every day of their lives.
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